So far we have discussed the transport of water and minerals in plants. Now let us consider how the products of metabolic processes, particularly photosynthesis, are moved from leaves, where they are formed, to other parts of the plant. This transport of soluble products of photosynthesis is called translocation and it occurs in the part of the vascular tissue known as phloem. Besides the products of photosynthesis, the phloem transports amino acids and other substances. These substances are especially delivered to the storage organs of roots, fruits and seeds and to growing organs. The translocation of food and other substances takes place in the sieve tubes with the help of adjacent companion cells both in upward and downward directions.
Unlike transport in xylem which can be largely explained by simple physical forces, the translocation in phloem is achieved by utilisingenergy. Material like sucrose is transferred into phloem tissue using energy from ATP. This increases the osmotic pressure of the tissue causing water to move into it. This pressure moves the material in the phloem to tissues which have less pressure. This allows the phloem to move material according to the plant’s needs. For example, in the spring, sugar stored in root or stem tissue would be transported to the buds which need energy to grow.
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